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The mission of the Quantitative Undergraduate Biology Education and Synthesis (QUBES) project is to improve learning opportunities for all students enrolled in undergraduate biology courses by reflecting the centrality of quantitative approaches in modern biology. The five major QUBES initiatives include: 1) QUBES Consortium: coordinating the efforts and resources of disparate communities invested in promoting quantitative biology education; 2) QUBES Faculty Mentoring Networks: supporting faculty understanding and implementation of specific quantitative biology concepts and teaching approaches; 3) QUBES Hub: increasing the visibility, utility, and adoption of existing quantitative biology materials and the capacity for peer educator interaction; 4) QUBES Metrics: quantifying and tracking faculty contribution to quantitative biology education scholarship; and, 5) Implementation Research: studying and disseminating the features of QUBES that increase implementation success.